About the Authors

Edward E. Large

Affiliation School of Biology, Georgia Institute of Technology, Atlanta, Georgia, United States of America

Wen Xu

Affiliation School of Biology, Georgia Institute of Technology, Atlanta, Georgia, United States of America

Yuehui Zhao

Affiliation School of Biology, Georgia Institute of Technology, Atlanta, Georgia, United States of America

Shannon C. Brady

Affiliation Department of Molecular Biosciences, Northwestern University, Evanston, Illinois, United States of America

Lijiang Long

Affiliation School of Biology, Georgia Institute of Technology, Atlanta, Georgia, United States of America

Rebecca A. Butcher

Affiliation Department of Chemistry, University of Florida, Gainesville, Florida, United States of America

Erik C. Andersen

Affiliation Department of Molecular Biosciences, Northwestern University, Evanston, Illinois, United States of America

Patrick T. McGrath

patrick.mcgrath@biology.gatech.edu

Affiliation School of Biology, Georgia Institute of Technology, Atlanta, Georgia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EEL ECA PTM. Performed the experiments: EEL WX YZ SCB LL. Analyzed the data: EEL ECA SCB PTM. Contributed reagents/materials/analysis tools: RAB. Wrote the paper: EEL PTM.