About the Authors

Qianqian Yu

Affiliation The Key Laboratory of Plant Cell Engineering and Germplasm Innovation, Ministry of Education, College of Life Science, Shandong University, Jinan, People’s Republic of China

Huiyu Tian

Affiliation The Key Laboratory of Plant Cell Engineering and Germplasm Innovation, Ministry of Education, College of Life Science, Shandong University, Jinan, People’s Republic of China

Kun Yue

Affiliation The Key Laboratory of Plant Cell Engineering and Germplasm Innovation, Ministry of Education, College of Life Science, Shandong University, Jinan, People’s Republic of China

Jiajia Liu

Affiliation The Key Laboratory of Plant Cell Engineering and Germplasm Innovation, Ministry of Education, College of Life Science, Shandong University, Jinan, People’s Republic of China

Bing Zhang

Affiliation The Key Laboratory of Plant Cell Engineering and Germplasm Innovation, Ministry of Education, College of Life Science, Shandong University, Jinan, People’s Republic of China

Xugang Li

Affiliation School of Biological Science and Technology, University of Jinan, Jinan, People’s Republic of China

Zhaojun Ding

dingzhaojun@sdu.edu.cn

Affiliation The Key Laboratory of Plant Cell Engineering and Germplasm Innovation, Ministry of Education, College of Life Science, Shandong University, Jinan, People’s Republic of China

Competing Interests

The authors have delcared that no competing interets exist.

Author Contributions

Conceived and designed the experiments: ZD. Performed the experiments: QY HT JL BZ. Analyzed the data: QY HT JL BZ KY. Wrote the paper: ZD HT KY XL.