About the Authors

T. Niroshini Senaratne

Affiliation Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Eric F. Joyce

Current address: Department of Genetics, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Affiliation Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Son C. Nguyen

Current address: Department of Genetics, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Affiliation Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

C.-ting Wu

twu@genetics.med.harvard.edu

Affiliation Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TNS EFJ CtW. Performed the experiments: TNS SCN EFJ. Analyzed the data: TNS EFJ. Contributed reagents/materials/analysis tools: TNS EFJ. Wrote the paper: TNS EFJ CtW.