About the Authors

Jennifer Zieba

Affiliations Department of Human Genetics, David Geffen School of Medicine at the University of California at Los Angeles, Los Angeles, California, United States of America, Department of Orthopaedic Surgery, David Geffen School of Medicine at the University of California at Los Angeles, Los Angeles, California, United States of America

Kimberly Nicole Forlenza

Affiliation Department of Orthopaedic Surgery, David Geffen School of Medicine at the University of California at Los Angeles, Los Angeles, California, United States of America

Jagteshwar Singh Khatra

Affiliation Department of Orthopaedic Surgery, David Geffen School of Medicine at the University of California at Los Angeles, Los Angeles, California, United States of America

Anna Sarukhanov

Affiliation Department of Orthopaedic Surgery, David Geffen School of Medicine at the University of California at Los Angeles, Los Angeles, California, United States of America

Ivan Duran

Affiliation Department of Orthopaedic Surgery, David Geffen School of Medicine at the University of California at Los Angeles, Los Angeles, California, United States of America

Diana Rigueur

Affiliation Department of Molecular, Cell, and Developmental Biology, University of California at Los Angeles, Los Angeles, California, United States of America

Karen M. Lyons

Affiliations Department of Orthopaedic Surgery, David Geffen School of Medicine at the University of California at Los Angeles, Los Angeles, California, United States of America, Department of Molecular, Cell, and Developmental Biology, University of California at Los Angeles, Los Angeles, California, United States of America

Daniel H. Cohn

Affiliations Department of Orthopaedic Surgery, David Geffen School of Medicine at the University of California at Los Angeles, Los Angeles, California, United States of America, Department of Molecular, Cell, and Developmental Biology, University of California at Los Angeles, Los Angeles, California, United States of America

Amy E. Merrill

Affiliations Center for Craniofacial Molecular Biology, Ostrow School of Dentistry, University of Southern California, Los Angeles, California, United States of America, Department of Biochemistry and Molecular Biology, Keck School of Medicine, University of Southern California, Los Angeles, California, United States of America

Deborah Krakow

dkrakow@mednet.ucla.edu

Affiliations Department of Human Genetics, David Geffen School of Medicine at the University of California at Los Angeles, Los Angeles, California, United States of America, Department of Orthopaedic Surgery, David Geffen School of Medicine at the University of California at Los Angeles, Los Angeles, California, United States of America, Department of Obstetrics and Gynecology, David Geffen School of Medicine at the University of California at Los Angeles, Los Angeles, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DK JZ AEM KML DHC. Performed the experiments: JZ KNF JSK ID DR AEM AS. Analyzed the data: JZ AEM DK KML. Contributed reagents/materials/analysis tools: DK KML. Wrote the paper: DK JZ KML AEM DHC.