About the Authors

Jun Ding

jun.ding@nih.gov (JD); SchlessingerD@grc.nia.nih.gov (DS)

Affiliation Laboratory of Genetics, National Institute on Aging, NIH, Baltimore, Maryland

Carlo Sidore

Affiliations Department of Biostatistics and Center for Statistical Genetics, University of Michigan, Ann Arbor, Michigan, Istituto di Ricerca Genetica e Biomedica, Consiglio Nazionale delle Ricerche, Monserrato, Cagliari, Italy, Università degli Studi di Sassari, Sassari, Italy

Thomas J. Butler

Affiliation Laboratory of Genetics, National Institute on Aging, NIH, Baltimore, Maryland

Mary Kate Wing

Affiliation Department of Biostatistics and Center for Statistical Genetics, University of Michigan, Ann Arbor, Michigan

Yong Qian

Affiliation Laboratory of Genetics, National Institute on Aging, NIH, Baltimore, Maryland

Osorio Meirelles

Affiliation Laboratory of Genetics, National Institute on Aging, NIH, Baltimore, Maryland

Fabio Busonero

Affiliations Department of Biostatistics and Center for Statistical Genetics, University of Michigan, Ann Arbor, Michigan, Istituto di Ricerca Genetica e Biomedica, Consiglio Nazionale delle Ricerche, Monserrato, Cagliari, Italy

Lam C. Tsoi

Affiliation Department of Biostatistics and Center for Statistical Genetics, University of Michigan, Ann Arbor, Michigan

Andrea Maschio

Affiliations Department of Biostatistics and Center for Statistical Genetics, University of Michigan, Ann Arbor, Michigan, Istituto di Ricerca Genetica e Biomedica, Consiglio Nazionale delle Ricerche, Monserrato, Cagliari, Italy

Andrea Angius

Affiliation Istituto di Ricerca Genetica e Biomedica, Consiglio Nazionale delle Ricerche, Monserrato, Cagliari, Italy

Hyun Min Kang

Affiliation Department of Biostatistics and Center for Statistical Genetics, University of Michigan, Ann Arbor, Michigan

Ramaiah Nagaraja

Affiliation Laboratory of Genetics, National Institute on Aging, NIH, Baltimore, Maryland

Francesco Cucca

Affiliations Istituto di Ricerca Genetica e Biomedica, Consiglio Nazionale delle Ricerche, Monserrato, Cagliari, Italy, Università degli Studi di Sassari, Sassari, Italy

Gonçalo R. Abecasis

Affiliation Department of Biostatistics and Center for Statistical Genetics, University of Michigan, Ann Arbor, Michigan

David Schlessinger

jun.ding@nih.gov (JD); SchlessingerD@grc.nia.nih.gov (DS)

Affiliation Laboratory of Genetics, National Institute on Aging, NIH, Baltimore, Maryland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JD FC GRA DS. Performed the experiments: FB AM AA RN FC GRA. Analyzed the data: JD CS TJB YQ OM. Contributed reagents/materials/analysis tools: MKW LCT HMK. Wrote the paper: JD CS RN FC GRA DS.