About the Authors

Tia DiTommaso

Affiliation Department of Biochemistry and Molecular Biology, Monash University, Clayton, Melbourne, Australia

Lynelle K. Jones

Affiliation Department of Biochemistry and Molecular Biology, Monash University, Clayton, Melbourne, Australia

Denny L. Cottle

Affiliation Department of Biochemistry and Molecular Biology, Monash University, Clayton, Melbourne, Australia

The WTSI Mouse Genetics Program

Membership of the WTSI Mouse Genetics Program is provided in the Acknowledgments.

Affiliation Wellcome Trust Sanger Institute, Genome Campus, Hinxton, Cambridge, United Kingdom

Anna-Karin Gerdin

Affiliation Wellcome Trust Sanger Institute, Genome Campus, Hinxton, Cambridge, United Kingdom

Valerie E. Vancollie

Affiliation Wellcome Trust Sanger Institute, Genome Campus, Hinxton, Cambridge, United Kingdom

Fiona M. Watt

Affiliation Centre for Stem Cells and Regenerative Medicine King's College London, Guy's Hospital, London, United Kingdom

Ramiro Ramirez-Solis

Affiliation Wellcome Trust Sanger Institute, Genome Campus, Hinxton, Cambridge, United Kingdom

Allan Bradley

Affiliation Wellcome Trust Sanger Institute, Genome Campus, Hinxton, Cambridge, United Kingdom

Karen P. Steel

Affiliations Wellcome Trust Sanger Institute, Genome Campus, Hinxton, Cambridge, United Kingdom, Wolfson Centre for Age-Related Diseases, King's College London, Guy's Campus, London, United Kingdom

John P. Sundberg

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Jacqueline K. White

Affiliation Wellcome Trust Sanger Institute, Genome Campus, Hinxton, Cambridge, United Kingdom

Ian M. Smyth

ian.smyth@monash.edu

Affiliations Department of Biochemistry and Molecular Biology, Monash University, Clayton, Melbourne, Australia, Department of Anatomy and Developmental Biology, Monash University, Clayton, Melbourne, Australia

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TD DLC RRS KPS JKW IMS. Performed the experiments: TD LKJ AKG. Analyzed the data: TD DLC VEV KPS JPS JKW IMS. Contributed reagents/materials/analysis tools: FMW AB. Wrote the paper: TD DLC IMS.