About the Authors

Kurt Warnhoff

Affiliation Department of Developmental Biology, Washington University School of Medicine, St. Louis, Missouri, United States of America

John T. Murphy

Affiliation Department of Developmental Biology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Sandeep Kumar

Affiliation Department of Developmental Biology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Daniel L. Schneider

Affiliation Department of Developmental Biology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Michelle Peterson

Affiliation Department of Developmental Biology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Simon Hsu

Affiliation Department of Developmental Biology, Washington University School of Medicine, St. Louis, Missouri, United States of America

James Guthrie

Affiliation Research Reactor Center, University of Missouri, Columbia, Missouri, United States of America

J. David Robertson

Affiliations Research Reactor Center, University of Missouri, Columbia, Missouri, United States of America, Department of Chemistry, University of Missouri, Columbia, Missouri, United States of America

Kerry Kornfeld

kornfeld@wustl.edu

Affiliation Department of Developmental Biology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KW JTM SK JG JDR KK. Performed the experiments: KW JTM SK DLS MP SH JG JDR. Analyzed the data: KW JTM SK DLS MP SH JG JDR KK. Contributed reagents/materials/analysis tools: KW JTM DLS JG JDR KK. Wrote the paper: KW KK.