About the Authors

Eryn Wicklow

Contributed equally to this work with: Eryn Wicklow, Stephanie Blij

Affiliation Molecular, Cell, and Developmental Biology, University of California Santa Cruz, Santa Cruz, California, United States of America

Stephanie Blij

Contributed equally to this work with: Eryn Wicklow, Stephanie Blij

Affiliation Molecular, Cell, and Developmental Biology, University of California Santa Cruz, Santa Cruz, California, United States of America

Tristan Frum

Current address: Biochemistry and Molecular Biology, Michigan State University, East Lansing, Michigan, United States of America

Affiliation Molecular, Cell, and Developmental Biology, University of California Santa Cruz, Santa Cruz, California, United States of America

Yoshikazu Hirate

Current address: Center for Experimental Animals, Tokyo Medical and Dental University, Tokyo, Japan

Affiliation Institute of Molecular Embryology and Genetics, Kumamoto University, Chuo-ku, Kumamoto, Japan

Richard A. Lang

Affiliation Division of Pediatric Ophthalmology, Cincinnati Children's Hospital Medical Center, Cincinnati, Ohio, United States of America

Hiroshi Sasaki

Affiliation Institute of Molecular Embryology and Genetics, Kumamoto University, Chuo-ku, Kumamoto, Japan

Amy Ralston

aralston@msu.edu

Current address: Biochemistry and Molecular Biology, Michigan State University, East Lansing, Michigan, United States of America

Affiliation Molecular, Cell, and Developmental Biology, University of California Santa Cruz, Santa Cruz, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EW SB TF YH AR. Performed the experiments: EW SB TF YH. Analyzed the data: EW SB TF YH AR. Contributed reagents/materials/analysis tools: HS RAL AR. Wrote the paper: AR.