About the Authors

Sasha A. Langley

Affiliation Life Sciences Division, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America

Gary H. Karpen

Affiliations Life Sciences Division, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America, Department of Molecular and Cell Biology, University of California - Berkeley, Berkeley, California, United States of America

Charles H. Langley

chlangley@ucdavis.edu

Affiliation Department of Evolution and Ecology, University of California - Davis, Davis, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SAL GHK CHL. Performed the experiments: SAL CHL. Analyzed the data: SAL CHL. Contributed reagents/materials/analysis tools: SAL GHK CHL. Wrote the paper: SAL GHK CHL.