About the Authors

Zena T. Wolf

Affiliation Department of Population Health and Reproduction, School of Veterinary Medicine, University of California, Davis, Davis, California, United States of America

Elizabeth J. Leslie

Affiliation Department of Pediatrics, University of Iowa, Iowa City, Iowa, United States of America

Boaz Arzi

Affiliation Department of Surgical and Radiological Sciences, School of Veterinary Medicine, University of California, Davis, Davis, California, United States of America

Kartika Jayashankar

Affiliation Department of Population Health and Reproduction, School of Veterinary Medicine, University of California, Davis, Davis, California, United States of America

Nili Karmi

Affiliation Department of Population Health and Reproduction, School of Veterinary Medicine, University of California, Davis, Davis, California, United States of America

Zhonglin Jia

Affiliations Department of Pediatrics, University of Iowa, Iowa City, Iowa, United States of America, State Key Laboratory of Oral Diseases, West China Hospital of Stomatology, Sichuan University, Chengdu, China, Department of Cleft Lip and Palate Surgery, West China Hospital of Stomatology, Sichuan University, Chengdu, China

Douglas J. Rowland

Affiliation Center for Molecular and Genomic Imaging, University of California, Davis, Davis, California, United States of America

Amy Young

Affiliation Department of Animal Science, University of California, Davis, Davis, California, United States of America

Noa Safra

Affiliation Department of Population Health and Reproduction, School of Veterinary Medicine, University of California, Davis, Davis, California, United States of America

Saundra Sliskovic

Affiliation Department of Population Health and Reproduction, School of Veterinary Medicine, University of California, Davis, Davis, California, United States of America

Jeffrey C. Murray

Affiliation Department of Pediatrics, University of Iowa, Iowa City, Iowa, United States of America

Claire M. Wade

Affiliation Faculty of Veterinary Science, University of Sydney, Sydney, New South Wales, Australia

Danika L. Bannasch

dlbannasch@ucdavis.edu

Affiliation Department of Population Health and Reproduction, School of Veterinary Medicine, University of California, Davis, Davis, California, United States of America

Competing Interests

A DNA test for the DLX6 mutation in Nova Scotia Duck Tolling Retriever is commercially available through BDH Genetics, which is owned by a family member of DLB. This commercial venture had no influence on the interpretation of the results presented within the manuscript.

Author Contributions

Conceived and designed the experiments: ZTW BA EJL NK JCM CMW DLB. Performed the experiments: ZTW EJL DJR KJ NK ZJ AY SS. Analyzed the data: ZTW EJL BA NK NS KJ JCM CMW DLB. Contributed reagents/materials/analysis tools: JCM DLB. Wrote the paper: ZTW EJL BA KJ DLB.