About the Authors

Jianquan Chen

Affiliation Department of Orthopaedic Surgery, Washington University School of Medicine, St. Louis, Missouri, United States of America

Xiaolin Tu

Current address: Department of Anatomy and Cell Biology, Indiana University School of Medicine, Indianapolis, Indiana, United States of America.

Affiliation Department of Medicine, Washington University School of Medicine, St. Louis, Missouri, United States of America

Emel Esen

Affiliations Department of Orthopaedic Surgery, Washington University School of Medicine, St. Louis, Missouri, United States of America, Division of Biology and Biomedical Sciences, Washington University in St. Louis, St. Louis, Missouri, United States of America

Kyu Sang Joeng

Current address: Department of Molecular and Human Genetics, Baylor College of Medicine, Houston, Texas, United States of America.

Affiliations Department of Medicine, Washington University School of Medicine, St. Louis, Missouri, United States of America, Division of Biology and Biomedical Sciences, Washington University in St. Louis, St. Louis, Missouri, United States of America

Congxin Lin

Affiliation Department of Medicine, Washington University School of Medicine, St. Louis, Missouri, United States of America

Jeffrey M. Arbeit

Affiliation Department of Surgery, Washington University School of Medicine, St. Louis, Missouri, United States of America

Markus A. Rüegg

Affiliation Biozentrum, University of Basel, Basel, Switzerland

Michael N. Hall

Affiliation Biozentrum, University of Basel, Basel, Switzerland

Liang Ma

Affiliations Department of Medicine, Washington University School of Medicine, St. Louis, Missouri, United States of America, Division of Biology and Biomedical Sciences, Washington University in St. Louis, St. Louis, Missouri, United States of America, Department of Developmental Biology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Fanxin Long

longf@wudosis.wustl.edu

Affiliations Department of Orthopaedic Surgery, Washington University School of Medicine, St. Louis, Missouri, United States of America, Department of Medicine, Washington University School of Medicine, St. Louis, Missouri, United States of America, Division of Biology and Biomedical Sciences, Washington University in St. Louis, St. Louis, Missouri, United States of America, Department of Developmental Biology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FL. Performed the experiments: JC XT EE KSJ CL. Analyzed the data: JC XT FL. Contributed reagents/materials/analysis tools: JMA MAR MNH LM. Wrote the paper: JC FL.