About the Authors

Eric F. Joyce

Affiliation Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Nicholas Apostolopoulos

Affiliation Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Brian J. Beliveau

Affiliation Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

C. -ting Wu

twu@genetics.med.harvard.edu

Affiliation Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EFJ CtW. Performed the experiments: EFJ. Analyzed the data: EFJ. Contributed reagents/materials/analysis tools: EFJ NA BJB. Wrote the paper: EFJ CtW.