About the Authors

Daria V. Zhernakova

Affiliation University of Groningen, University Medical Center Groningen, Department of Genetics, Groningen, The Netherlands

Eleonora de Klerk

Contributed equally to this work with: Eleonora de Klerk, Harm-Jan Westra

Affiliation Center for Human and Clinical Genetics, Leiden University Medical Center, Leiden, The Netherlands

Harm-Jan Westra

Contributed equally to this work with: Eleonora de Klerk, Harm-Jan Westra

Affiliation University of Groningen, University Medical Center Groningen, Department of Genetics, Groningen, The Netherlands

Anastasios Mastrokolias

Affiliation Center for Human and Clinical Genetics, Leiden University Medical Center, Leiden, The Netherlands

Shoaib Amini

Affiliation Center for Human and Clinical Genetics, Leiden University Medical Center, Leiden, The Netherlands

Yavuz Ariyurek

Affiliations Center for Human and Clinical Genetics, Leiden University Medical Center, Leiden, The Netherlands, Leiden Genome Technology Center, Leiden, The Netherlands

Rick Jansen

Affiliation Department of Psychiatry, The Netherlands Study of Depression and Anxiety, VU University Medical Center, Amsterdam, The Netherlands

Brenda W. Penninx

Affiliation Department of Psychiatry, The Netherlands Study of Depression and Anxiety, VU University Medical Center, Amsterdam, The Netherlands

Jouke J. Hottenga

Affiliation Department of Biological Psychology, Netherlands Twin Registry, VU University, Amsterdam, The Netherlands

Gonneke Willemsen

Affiliation Department of Biological Psychology, Netherlands Twin Registry, VU University, Amsterdam, The Netherlands

Eco J. de Geus

Affiliation Department of Biological Psychology, Netherlands Twin Registry, VU University, Amsterdam, The Netherlands

Dorret I. Boomsma

Affiliation Department of Biological Psychology, Netherlands Twin Registry, VU University, Amsterdam, The Netherlands

Jan H. Veldink

Affiliation Department of Neurology, Rudolf Magnus Institute of Neuroscience, University Medical Centre Utrecht, Utrecht, The Netherlands

Leonard H. van den Berg

Affiliation Department of Neurology, Rudolf Magnus Institute of Neuroscience, University Medical Centre Utrecht, Utrecht, The Netherlands

Cisca Wijmenga

Affiliation University of Groningen, University Medical Center Groningen, Department of Genetics, Groningen, The Netherlands

Johan T. den Dunnen

Affiliations Center for Human and Clinical Genetics, Leiden University Medical Center, Leiden, The Netherlands, Leiden Genome Technology Center, Leiden, The Netherlands

Gert-Jan B. van Ommen

Affiliation Center for Human and Clinical Genetics, Leiden University Medical Center, Leiden, The Netherlands

Peter A. C. 't Hoen

These authors also contributed equally to this work.

Affiliation Center for Human and Clinical Genetics, Leiden University Medical Center, Leiden, The Netherlands

Lude Franke

lude@ludesign.nl

These authors also contributed equally to this work.

Affiliation University of Groningen, University Medical Center Groningen, Department of Genetics, Groningen, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PACtH LF GJBvO JTdD. Performed the experiments: PACtH YA AM. Analyzed the data: DVZ EdK PACtH HJW SA. Contributed reagents/materials/analysis tools: PACtH HJW RJ BWP JJH GW EJdG DIB JHV LHvdB CW. Wrote the paper: DVZ EdK PACtH LF.