About the Authors

Mark R. Davies

Current address: The Wellcome Trust Sanger Institute, The Wellcome Trust Genome Campus, Hinxton, Cambridgeshire, United Kingdom

Affiliation Centre for Immunology and Infection, Hull York Medical School and the Department of Biology, University of York, York, United Kingdom

Sarah E. Broadbent

Current address: Department of Chemical Engineering and Biotechnology, University of Cambridge, Cambridge, United Kingdom

Affiliation Centre for Immunology and Infection, Hull York Medical School and the Department of Biology, University of York, York, United Kingdom

Simon R. Harris

Affiliation The Wellcome Trust Sanger Institute, The Wellcome Trust Genome Campus, Hinxton, Cambridgeshire, United Kingdom

Nicholas R. Thomson

Affiliation The Wellcome Trust Sanger Institute, The Wellcome Trust Genome Campus, Hinxton, Cambridgeshire, United Kingdom

Marjan W. van der Woude

marjan.vanderwoude@york.ac.uk

Affiliation Centre for Immunology and Infection, Hull York Medical School and the Department of Biology, University of York, York, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MRD SEB SRH NRT MWvdW. Performed the experiments: MRD SEB SRH. Analyzed the data: MRD SEB SRH NRT MWvdW. Wrote the paper: MRD SEB NRT MWvdW.