About the Authors

Elizabeth X. Kwan

Affiliation Clinical Research Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Eric J. Foss

Affiliation Clinical Research Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Scott Tsuchiyama

Affiliation The Buck Institute for Research on Aging, Novato, California, United States of America

Gina M. Alvino

Affiliation Department of Genome Sciences, University of Washington, Seattle, Washington, United States of America

Leonid Kruglyak

Affiliation Howard Hughes Medical Institute, Lewis-Sigler Institute for Integrative Genomics, and Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

Matt Kaeberlein

Affiliation Department of Pathology, University of Washington, Seattle, Washington, United States of America

M. K. Raghuraman

Affiliation Department of Genome Sciences, University of Washington, Seattle, Washington, United States of America

Bonita J. Brewer

Affiliation Department of Genome Sciences, University of Washington, Seattle, Washington, United States of America

Brian K. Kennedy

Affiliation The Buck Institute for Research on Aging, Novato, California, United States of America

Antonio Bedalov

abedalov@fhcrc.org

Affiliation Clinical Research Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EXK EJF MKR BJB BKK AB. Performed the experiments: EXK ST GMA MKR BJB. Analyzed the data: EXK EJF MK MKR BJB BKK AB. Contributed reagents/materials/analysis tools: LK. Wrote the paper: EXK EJF MKR BJB BKK AB.