About the Authors

Marc Haber

Affiliations Institut de Biologia Evolutiva (CSIC–UPF), Departament de Ciències de la Salut i de la Vida, Universitat Pompeu Fabra, Barcelona, Spain, The Lebanese American University, Chouran, Beirut, Lebanon

Dominique Gauguier

Affiliations The Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom, INSERM UMRS872, Centre de Recherche des Cordeliers, Paris, France

Sonia Youhanna

Affiliation The Lebanese American University, Chouran, Beirut, Lebanon

Nick Patterson

Affiliation Broad Institute of Harvard and MIT, Cambridge, Massachusetts, United States of America

Priya Moorjani

Affiliation Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Laura R. Botigué

Affiliation Institut de Biologia Evolutiva (CSIC–UPF), Departament de Ciències de la Salut i de la Vida, Universitat Pompeu Fabra, Barcelona, Spain

Daniel E. Platt

Affiliation Bioinformatics and Pattern Discovery, IBM T. J. Watson Research Centre, Yorktown Heights, New York, United States of America

Elizabeth Matisoo-Smith

Affiliation Allan Wilson Centre for Molecular Ecology and Evolution and Department of Anatomy, University of Otago, Dunedin, New Zealand

David F. Soria-Hernanz

Affiliation The Genographic Project, National Geographic Society, Washington, D.C., United States of America

R. Spencer Wells

Affiliation The Genographic Project, National Geographic Society, Washington, D.C., United States of America

Jaume Bertranpetit

Affiliation Institut de Biologia Evolutiva (CSIC–UPF), Departament de Ciències de la Salut i de la Vida, Universitat Pompeu Fabra, Barcelona, Spain

Chris Tyler-Smith

Affiliation The Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, United Kingdom

David Comas

Contributed equally to this work with: David Comas, Pierre A. Zalloua

Affiliation Institut de Biologia Evolutiva (CSIC–UPF), Departament de Ciències de la Salut i de la Vida, Universitat Pompeu Fabra, Barcelona, Spain

Pierre A. Zalloua

Contributed equally to this work with: David Comas, Pierre A. Zalloua

pierre.zalloua@lau.edu.lb

Affiliations The Lebanese American University, Chouran, Beirut, Lebanon, Harvard School of Public Health, Boston, Massachusetts, United States of America

Competing Interests

DEP is an employee of IBM. There are no patents or products in development or marketed products to declare. This does not alter the author's adherence to all the PLOS Genetics policies on sharing data and materials. All other authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MH DC PAZ. Performed the experiments: MH PM. Analyzed the data: MH NP PM DFS-H CT-S DC PAZ. Contributed reagents/materials/analysis tools: DG SY LRB DEP EM-S RSW JB. Wrote the paper: MH DC PAZ.