About the Authors
- Noah Zaitlen
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* E-mail: nzaitlen@hsph.harvard.edu (N Zaitlen); aprice@hsph.harvard.edu (AL Price)
Affiliations Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America, Department of Biostatistics, Harvard School of Public Health, Boston, Massachusetts, United States of America, Broad Institute of Harvard and Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America, Program in Molecular and Genetic Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America
- Sara Lindström
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Affiliations Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America, Program in Molecular and Genetic Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America
- Bogdan Pasaniuc
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Affiliations Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America, Department of Biostatistics, Harvard School of Public Health, Boston, Massachusetts, United States of America, Broad Institute of Harvard and Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America, Program in Molecular and Genetic Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America
- Marilyn Cornelis
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Affiliation Channing Laboratory, Department of Medicine, Brigham and Women's Hospital and Harvard Medical School, Boston, Massachusetts, United States of America
- Giulio Genovese
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Affiliation Division of Nephrology, Department of Medicine, Beth Israel Deaconess Medical Center and Harvard Medical School, Boston, Massachusetts, United States of America
- Samuela Pollack
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Affiliations Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America, Department of Biostatistics, Harvard School of Public Health, Boston, Massachusetts, United States of America, Broad Institute of Harvard and Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America, Program in Molecular and Genetic Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America
- Anne Barton
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Affiliation Arthritis Research UK Epidemiology Unit, Manchester Academic Health Science Centre, The University of Manchester, Manchester, United Kingdom
- Heike Bickeböller
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Affiliation Department of Genetic Epidemiology, University Medical Centre, University of Göttingen, Göttingen, Germany
- Donald W. Bowden
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Affiliation Center for Human Genomics, Wake Forest School of Medicine, Winston-Salem, North Carolina, United States of America
- Steve Eyre
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Affiliation Arthritis Research UK Epidemiology Unit, Manchester Academic Health Science Centre, The University of Manchester, Manchester, United Kingdom
- Barry I. Freedman
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Affiliation Department of Internal Medicine/Section on Nephrology, Wake Forest School of Medicine, Winston-Salem, North Carolina, United States of America
- David J. Friedman
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Affiliation Division of Nephrology, Department of Medicine, Beth Israel Deaconess Medical Center and Harvard Medical School, Boston, Massachusetts, United States of America
- John K. Field
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Affiliation Roy Castle Lung Cancer Research Programme, University of Liverpool, Liverpool, United Kingdom
- Leif Groop
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Affiliation Department of Clinical Sciences, Diabetes and Endocrinology Research Unit, Scania University Hospital, Lund University, Malmö, Sweden
- Aage Haugen
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Affiliation Section for Toxicology, National Institute of Occupational Health, Oslo, Norway
- Joachim Heinrich
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Affiliation Institute of Epidemiology, German Research Centre for Environmental Health, Neuherberg, Germany
- Brian E. Henderson
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Affiliation Department of Preventive Medicine, University of Southern California Keck School of Medicine, Los Angeles, California, United States of America
- Pamela J. Hicks
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Affiliation Department of Biochemistry, Wake Forest University School of Medicine, Winston-Salem, North Carolina, United States of America
- Lynne J. Hocking
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Affiliation Musculoskeletal Research Programme, Division of Applied Medicine, University of Aberdeen, Aberdeen, United Kingdom
- Laurence N. Kolonel
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Affiliation Epidemiology Program, Cancer Research Center, University of Hawaii, Honolulu, Hawaii, United States of America
- Maria Teresa Landi
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Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, Bethesda, Maryland, United States of America
- Carl D. Langefeld
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Affiliation Division of Public Health Sciences, Wake Forest University School of Medicine, Winston-Salem, North Carolina, United States of America
- Loic Le Marchand
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Affiliation Epidemiology Program, Cancer Research Center, University of Hawaii, Honolulu, Hawaii, United States of America
- Michael Meister
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Affiliations Thoraxklinik am Universitätsklinikum, Heidelberg, Germany, Translational Lung Research Centre Heidelberg (TLRC-H), German Center for Lung Research, Heidelberg, Germany
- Ann W. Morgan
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Affiliation NIHR–Leeds Musculoskeletal Biomedical Research Unit, Leeds, United Kingdom
- Olaide Y. Raji
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Affiliation Roy Castle Lung Cancer Research Programme, University of Liverpool, Liverpool, United Kingdom
- Angela Risch
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Affiliations Translational Lung Research Centre Heidelberg (TLRC-H), German Center for Lung Research, Heidelberg, Germany, DKFZ–German Cancer Research Center, Heidelberg, Germany
- Albert Rosenberger
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Affiliation Department of Genetic Epidemiology, University Medical Centre, University of Göttingen, Göttingen, Germany
- David Scherf
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Affiliation DKFZ–German Cancer Research Center, Heidelberg, Germany
- Sophia Steer
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Affiliation King's College Hospital National Health Service Foundation Trust, London, United Kingdom
- Martin Walshaw
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Affiliation Liverpool Heart and Chest Hospital, Liverpool, United Kingdom
- Kevin M. Waters
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Affiliation Department of Preventive Medicine, University of Southern California Keck School of Medicine, Los Angeles, California, United States of America
- Anthony G. Wilson
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Affiliation Deptartment of Infection and Immunity, University of Sheffield, Sheffield, United Kingdom
- Paul Wordsworth
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Affiliation NIHR Oxford Musculoskeletal Biomedical Research Unit, Nuffield Orthopaedic Centre, Oxford, United Kingdom
- Shanbeh Zienolddiny
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Affiliation Section for Toxicology, National Institute of Occupational Health, Oslo, Norway
- Eric Tchetgen Tchetgen
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Affiliations Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America, Department of Biostatistics, Harvard School of Public Health, Boston, Massachusetts, United States of America
- Christopher Haiman
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Affiliation Department of Preventive Medicine, University of Southern California Keck School of Medicine, Los Angeles, California, United States of America
- David J. Hunter
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Affiliations Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America, Broad Institute of Harvard and Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America, Program in Molecular and Genetic Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America, Channing Laboratory, Department of Medicine, Brigham and Women's Hospital and Harvard Medical School, Boston, Massachusetts, United States of America
- Robert M. Plenge
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Affiliations Broad Institute of Harvard and Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America, Division of Rheumatology, Immunology, and Allergy and Division of Genetics, Brigham and Women's Hospital, Boston, Massachusetts, United States of America
- Jane Worthington
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Affiliation Arthritis Research UK Epidemiology Unit, Manchester Academic Health Science Centre, The University of Manchester, Manchester, United Kingdom
- David C. Christiani
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Affiliations Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America, Department of Environmental Health, Harvard School of Public Health, Boston, Massachusetts, United States of America
- Debra A. Schaumberg
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Affiliations Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America, Division of Preventive Medicine, Department of Medicine, Brigham and Women's Hospital, Boston, Harvard Medical School, Massachusetts, United States of America, Schepens Eye Research Institute, Department of Ophthalmology, Harvard Medical School, Boston, Massachusetts, United States of America
- Daniel I. Chasman
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Affiliation Schepens Eye Research Institute, Department of Ophthalmology, Harvard Medical School, Boston, Massachusetts, United States of America
- David Altshuler
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Affiliations Broad Institute of Harvard and Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America, Center for Human Genetic Research, Department of Molecular Biology and Diabetes Unit, Massachusetts General Hospital, Boston, Massachusetts, United States of America, Departments of Genetics and Medicine, Harvard Medical School, Boston, Massachusetts, United States of America
- Benjamin Voight
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Affiliations Broad Institute of Harvard and Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America, Center for Human Genetic Research, Department of Molecular Biology and Diabetes Unit, Massachusetts General Hospital, Boston, Massachusetts, United States of America, Departments of Genetics and Medicine, Harvard Medical School, Boston, Massachusetts, United States of America
- Peter Kraft
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Affiliations Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America, Department of Biostatistics, Harvard School of Public Health, Boston, Massachusetts, United States of America, Broad Institute of Harvard and Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America, Program in Molecular and Genetic Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America
- Nick Patterson
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Affiliation Broad Institute of Harvard and Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America
- Alkes L. Price
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* E-mail: nzaitlen@hsph.harvard.edu (N Zaitlen); aprice@hsph.harvard.edu (AL Price)
Affiliations Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America, Department of Biostatistics, Harvard School of Public Health, Boston, Massachusetts, United States of America, Broad Institute of Harvard and Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America, Program in Molecular and Genetic Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America
Competing Interests
The authors have declared that no competing interests exist.
Author Contributions
Conceived and designed the experiments: N Zaitlen, N Patterson, AL Price, P Kraft, E Tchetgen Tchetgen. Performed the experiments: N Zaitlen, AL Price, S Pollack. Analyzed the data: N Zaitlen, AL Price, B Pasaniuc, P Kraft. Contributed reagents/materials/analysis tools: M Cornelis, G Genovese, A Barton, H Bickeböller, DW Bowden, S Eyre, BI Freedman, DJ Friedman, JK Field, L Groop, A Haugen, J Heinrich, BE Henderson, PJ Hicks, LJ Hocking, LN Kolonel, MT Landi, CD Langefeld, L Le Marchand, M Meister, AW Morgan, OY Raji, A Rosenberger, A Risch, D Scherf, S Steer, M Walshaw, KM Waters, AG Wilson, P Wordsworth, S Zienolddiny, C Haiman, DJ Hunter, RM Plenge, J Worthington, DC Christiani, DA Schaumberg, DI Chasman, D Altshuler, B Voight. Wrote the paper: N Zaitlen, S Lindström, AL Price.