About the Authors

Jennifer S. Yokoyama

Contributed equally to this work with: Jennifer S. Yokoyama, Ernest T. Lam

Affiliation Department of Psychiatry and Institute for Human Genetics, University of California San Francisco, San Francisco, California, United States of America

Ernest T. Lam

Contributed equally to this work with: Jennifer S. Yokoyama, Ernest T. Lam

Affiliation Institute for Human Genetics, University of California San Francisco, San Francisco, California, United States of America

Alison L. Ruhe

Affiliations Neurogenomics Division, The Translational Genomics Research Institute, Phoenix, Arizona, United States of America, Veterinary Genetics Laboratory, University of California Davis, Davis, California, United States of America

Carolyn A. Erdman

Affiliation Department of Psychiatry and Institute for Human Genetics, University of California San Francisco, San Francisco, California, United States of America

Kathryn R. Robertson

Affiliation Veterinary Genetics Laboratory, University of California Davis, Davis, California, United States of America

Aubrey A. Webb

Affiliations CullenWebb Animal Neurology and Ophthalmology Centre, Riverview, New Brunswick, Canada, Department of Clinical Neuroscience, Faculty of Medicine, University of Calgary, Calgary, Alberta, Canada

D. Colette Williams

Affiliation School of Veterinary Medicine, University of California Davis, Davis, California, United States of America

Melanie L. Chang

Affiliation Department of Anthropology, University of Oregon, Eugene, Oregon, United States of America

Marjo K. Hytönen

Affiliation Department of Veterinary Biosciences and Research Programs Unit, Molecular Medicine, University of Helsinki and Folkhälsan Research Center, Helsinki, Finland

Hannes Lohi

Affiliation Department of Veterinary Biosciences and Research Programs Unit, Molecular Medicine, University of Helsinki and Folkhälsan Research Center, Helsinki, Finland

Steven P. Hamilton

Affiliation Department of Psychiatry and Institute for Human Genetics, University of California San Francisco, San Francisco, California, United States of America

Mark W. Neff

mark.neff@vai.org

Affiliations Neurogenomics Division, The Translational Genomics Research Institute, Phoenix, Arizona, United States of America, The Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JSY ETL SPH MWN. Performed the experiments: JSY ETL ALR CAE KRR MKH. Analyzed the data: JSY ETL. Contributed reagents/materials/analysis tools: JSY ETL ALR CAE KRR AAW DCW MLC MKH HL SPH MWN. Wrote the paper: JSY ETL CAE AAW MLC SPH MWN. Critical reading and discussion of manuscript: AAW DCW HL ALR.