About the Authors

Ripla Arora

Affiliation Department of Genetics and Development, Columbia University Medical Center, New York, New York, United States of America

Ross J. Metzger

Affiliation Department of Anatomy, School of Medicine, University of California San Francisco, San Francisco, California, United States of America

Virginia E. Papaioannou

vep1@columbia.edu

Affiliation Department of Genetics and Development, Columbia University Medical Center, New York, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RA VEP. Performed the experiments: RA VEP. Analyzed the data: RA VEP RJM. Contributed reagents/materials/analysis tools: RA VEP. Wrote the paper: RA VEP RJM.