About the Authors

Madapura M. Pradeepa

Affiliation MRC Human Genetics Unit, MRC Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, United Kingdom

Heidi G. Sutherland

Affiliation MRC Human Genetics Unit, MRC Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, United Kingdom

Jernej Ule

Affiliation MRC Laboratory of Molecular Biology, Cambridge, United Kingdom

Graeme R. Grimes

Affiliation MRC Human Genetics Unit, MRC Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, United Kingdom

Wendy A. Bickmore

W.Bickmore@hgu.mrc.ac.uk

Affiliation MRC Human Genetics Unit, MRC Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MMP WAB. Performed the experiments: MMP HGS JU. Analyzed the data: MMP JU GRG. Wrote the paper: MMP WAB.