About the Authors

Joseph P. Jarvis

Affiliation Department of Genetics, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Laura B. Scheinfeldt

Contributed equally to this work with: Laura B. Scheinfeldt, Sameer Soi, Charla Lambert, Larsson Omberg

Affiliation Department of Genetics, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Sameer Soi

Contributed equally to this work with: Laura B. Scheinfeldt, Sameer Soi, Charla Lambert, Larsson Omberg

Affiliation Department of Genetics, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Charla Lambert

Contributed equally to this work with: Laura B. Scheinfeldt, Sameer Soi, Charla Lambert, Larsson Omberg

Affiliation Department of Genetics, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Larsson Omberg

Contributed equally to this work with: Laura B. Scheinfeldt, Sameer Soi, Charla Lambert, Larsson Omberg

Affiliation Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America

Bart Ferwerda

Affiliation Department of Genetics, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Alain Froment

Affiliation UMR 208, IRD-MNHN, Musée de l'Homme, Paris, France

Jean-Marie Bodo

Affiliation Ministere de la Recherche Scientifique et de l'Innovation, Yaounde, Cameroon

William Beggs

Affiliation Department of Genetics, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Gabriel Hoffman

Affiliation Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America

Jason Mezey

Affiliations Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America, Department of Genetic Medicine, Weill Cornell Medical College, New York, New York, United States of America

Sarah A. Tishkoff

tishkoff@mail.med.upenn.edu

Affiliations Department of Genetics, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America, Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JPJ JM SAT. Performed the experiments: WB BF. Analyzed the data: JPJ LBS SS CL LO BF GH JM SAT. Contributed reagents/materials/analysis tools: AF J-MB GH LO JM. Wrote the paper: JPJ SAT.