About the Authors

Michael J. Guertin

Contributed equally to this work with: Michael J. Guertin, André L. Martins

Affiliation Department of Molecular Biology and Genetics, Cornell University, Ithaca, New York, United States of America

André L. Martins

Contributed equally to this work with: Michael J. Guertin, André L. Martins

Affiliation Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America

Adam Siepel

Affiliation Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America

John T. Lis

jtl10@cornell.edu

Affiliation Department of Molecular Biology and Genetics, Cornell University, Ithaca, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MJG JTL. Performed the experiments: MJG. Analyzed the data: MJG ALM. Wrote the paper: MJG ALM AS JTL. Developed the statistical models: ALM AS.