About the Authors

Kimberly Larson

Contributed equally to this work with: Kimberly Larson, Shian-Jang Yan, Amy Tsurumi, Jacqueline Liu

Affiliation Department of Biomedical Genetics, University of Rochester Medical Center, Rochester, New York, United States of America

Shian-Jang Yan

Contributed equally to this work with: Kimberly Larson, Shian-Jang Yan, Amy Tsurumi, Jacqueline Liu

Affiliations Department of Biomedical Genetics, University of Rochester Medical Center, Rochester, New York, United States of America, Department of Medicine, University of California San Diego, La Jolla, California, United States of America

Amy Tsurumi

Contributed equally to this work with: Kimberly Larson, Shian-Jang Yan, Amy Tsurumi, Jacqueline Liu

Affiliation Department of Biomedical Genetics, University of Rochester Medical Center, Rochester, New York, United States of America

Jacqueline Liu

Contributed equally to this work with: Kimberly Larson, Shian-Jang Yan, Amy Tsurumi, Jacqueline Liu

Affiliation Department of Biomedical Genetics, University of Rochester Medical Center, Rochester, New York, United States of America

Jun Zhou

Affiliation Department of Biology, University of Rochester, Rochester, New York, United States of America

Kriti Gaur

Affiliation Department of Biomedical Genetics, University of Rochester Medical Center, Rochester, New York, United States of America

Dongdong Guo

Affiliation Department of Biomedical Genetics, University of Rochester Medical Center, Rochester, New York, United States of America

Thomas H. Eickbush

Affiliation Department of Biology, University of Rochester, Rochester, New York, United States of America

Willis X. Li

willisli@ucsd.edu

Affiliations Department of Biomedical Genetics, University of Rochester Medical Center, Rochester, New York, United States of America, Department of Medicine, University of California San Diego, La Jolla, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WXL. Performed the experiments: KL S-JY AT JL JZ KG DG. Analyzed the data: KL S-JY AT JL JZ THE WXL. Wrote the paper: WXL.