About the Authors

Brenna M. Henn

Contributed equally to this work with: Brenna M. Henn, Laura R. Botigué

Affiliation Department of Genetics, Stanford University, Stanford, California, United States of America

Laura R. Botigué

Contributed equally to this work with: Brenna M. Henn, Laura R. Botigué

Affiliation Institute of Evolutionary Biology (CSIC-UPF), Universitat Pompeu Fabra, Barcelona, Spain

Simon Gravel

Affiliation Department of Genetics, Stanford University, Stanford, California, United States of America

Wei Wang

Affiliation Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America

Abra Brisbin

Affiliation Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America

Jake K. Byrnes

Affiliation Department of Genetics, Stanford University, Stanford, California, United States of America

Karima Fadhlaoui-Zid

Affiliation Laboratory of Genetics, Immunology, and Human Pathology, University Tunis El Manar, Tunis, Tunisia

Pierre A. Zalloua

Affiliation The Lebanese American University, Chouran, Beirut, Lebanon

Andres Moreno-Estrada

Affiliation Department of Genetics, Stanford University, Stanford, California, United States of America

Jaume Bertranpetit

Affiliation Institute of Evolutionary Biology (CSIC-UPF), Universitat Pompeu Fabra, Barcelona, Spain

Carlos D. Bustamante

david.comas@upf.edu (DC); cdbustam@stanford.edu (CDB)

These authors were joint senior authors on this work.

Affiliation Department of Genetics, Stanford University, Stanford, California, United States of America

David Comas

david.comas@upf.edu (DC); cdbustam@stanford.edu (CDB)

These authors were joint senior authors on this work.

Affiliation Institute of Evolutionary Biology (CSIC-UPF), Universitat Pompeu Fabra, Barcelona, Spain

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BMH SG JB CDB DC. Performed the experiments: BMH LRB. Analyzed the data: BMH LRB SG WW AM-E. Contributed reagents/materials/analysis tools: AB JKB KF-Z PAZ JB DC. Wrote the paper: BMH LRB SG CDB DC.