About the Authors

Laura Y. Fox-Clipsham

Affiliation Animal Health Trust, Newmarket, United Kingdom

Stuart D. Carter

Affiliation Department of Infection Biology, School of Veterinary Science, University of Liverpool, Liverpool, United Kingdom

Ian Goodhead

Affiliation Centre for Genomic Research, Institute of Integrative Biology, University of Liverpool, Liverpool, United Kingdom

Neil Hall

Affiliation Centre for Genomic Research, Institute of Integrative Biology, University of Liverpool, Liverpool, United Kingdom

Derek C. Knottenbelt

Affiliation Department of Veterinary Clinical Science, Equine Hospital, University of Liverpool, Liverpool, United Kingdom

Paul D. F. May

Affiliation Townhead Veterinary Centre, Townhead Farm, Penrith, United Kingdom

William E. Ollier

Affiliation Centre for Integrated Genomic Medical Research, University of Manchester, Manchester, United Kingdom

June E. Swinburne

june.swinburne@aht.org.uk

Affiliation Animal Health Trust, Newmarket, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SDC WEO JES. Performed the experiments: LYF-C IG JES. Analyzed the data: LYF-C IG JES. Contributed reagents/materials/analysis tools: NH DCK PDFM. Wrote the paper: JES.