About the Authors

Karin E. Smedby

Contributed equally to this work with: Karin E. Smedby, Jia Nee Foo

Affiliation Department of Medicine, Clinical Epidemiology Unit, Karolinska Institutet, Stockholm, Sweden

Jia Nee Foo

Contributed equally to this work with: Karin E. Smedby, Jia Nee Foo

Affiliation Human Genetics, Genome Institute of Singapore, A*STAR, Singapore, Singapore

Christine F. Skibola

‡ These authors also contributed equally to this work.

Affiliation Division of Environmental Health Sciences, School of Public Health, University of California Berkeley, Berkeley, California, United States of America

Hatef Darabi

‡ These authors also contributed equally to this work.

Affiliation Department of Medical Epidemiology and Biostatistics, Karolinska Institutet, Stockholm, Sweden

Lucia Conde

Affiliation Division of Environmental Health Sciences, School of Public Health, University of California Berkeley, Berkeley, California, United States of America

Henrik Hjalgrim

Affiliation Department of Epidemiology Research, Statens Serum Institut, Copenhagen, Denmark

Vikrant Kumar

Affiliation Human Genetics, Genome Institute of Singapore, A*STAR, Singapore, Singapore

Ellen T. Chang

Affiliations Cancer Prevention Institute of California, Fremont, California, United States of America, Division of Epidemiology, Department of Health Research and Policy, Stanford University School of Medicine, Stanford, California, United States of America

Nathaniel Rothman

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, Bethesda, Maryland, United States of America

James R. Cerhan

Affiliation College of Medicine, Mayo Clinic, Rochester, Minnesota, United States of America

Angela R. Brooks-Wilson

Affiliations British Columbia Cancer Research Center, British Columbia Cancer Agency, Vancouver, Canada, Department of Biomedical Physiology and Kinesiology, Simon Fraser University, Burnaby, Canada

Emil Rehnberg

Affiliation Department of Medical Epidemiology and Biostatistics, Karolinska Institutet, Stockholm, Sweden

Ishak D. Irwan

Affiliation Human Genetics, Genome Institute of Singapore, A*STAR, Singapore, Singapore

Lars P. Ryder

Affiliation Department of Clinical Immunology, University Hospital of Copenhagen, Copenhagen, Denmark

Peter N. Brown

Affiliation Department of Haematology, Copenhagen University Hospital, Copenhagen, Denmark

Paige M. Bracci

Affiliation Department of Epidemiology and Biostatistics, University of California San Francisco, San Francisco, California, United States of America

Luz Agana

Affiliation Division of Environmental Health Sciences, School of Public Health, University of California Berkeley, Berkeley, California, United States of America

Jacques Riby

Affiliation Division of Environmental Health Sciences, School of Public Health, University of California Berkeley, Berkeley, California, United States of America

Wendy Cozen

Affiliation Department of Preventive Medicine, Keck School of Medicine, University of Southern California, Los Angeles, California, United States of America

Scott Davis

Affiliations Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, University of Washington, Seattle, Washington, United States of America

Patricia Hartge

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Lindsay M. Morton

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Richard K. Severson

Affiliations Department of Family Medicine and Public Health Sciences, Wayne State University, Detroit, Michigan, United States of America, Karmanos Cancer Institute, Detroit, Michigan, United States of America

Sophia S. Wang

Affiliations Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, Bethesda, Maryland, United States of America, Division of Etiology, Beckman Research Institute and the City of Hope, Duarte, California, United States of America

Susan L. Slager

Affiliation College of Medicine, Mayo Clinic, Rochester, Minnesota, United States of America

Zachary S. Fredericksen

Affiliation College of Medicine, Mayo Clinic, Rochester, Minnesota, United States of America

Anne J. Novak

Affiliation College of Medicine, Mayo Clinic, Rochester, Minnesota, United States of America

Neil E. Kay

Affiliation College of Medicine, Mayo Clinic, Rochester, Minnesota, United States of America

Thomas M. Habermann

Affiliation College of Medicine, Mayo Clinic, Rochester, Minnesota, United States of America

Bruce Armstrong

Affiliation Sydney School of Public Health, The University of Sydney, Sydney, Australia

Anne Kricker

Affiliation Sydney School of Public Health, The University of Sydney, Sydney, Australia

Sam Milliken

Affiliation Department of Haematology, St. Vincent's Hospital, Sydney, Australia

Mark P. Purdue

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Claire M. Vajdic

Affiliation University of New South Wales Cancer Research Center, Prince of Wales Clinical School, Sydney, Australia

Peter Boyle

Affiliation International Prevention Research Institute, Lyon, France

Qing Lan

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Shelia H. Zahm

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Yawei Zhang

Affiliation School of Public Health, Yale University, New Haven, Connecticut, United States of America

Tongzhang Zheng

Affiliation School of Public Health, Yale University, New Haven, Connecticut, United States of America

Stephen Leach

Affiliation Canada's Michael Smith Genome Sciences Centre, British Columbia Cancer Agency, Vancouver, Canada

John J. Spinelli

Affiliations British Columbia Cancer Research Center, British Columbia Cancer Agency, Vancouver, Canada, School of Population and Public Health, University of British Columbia, Vancouver, Canada

Martyn T. Smith

Affiliation Division of Environmental Health Sciences, School of Public Health, University of California Berkeley, Berkeley, California, United States of America

Stephen J. Chanock

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Leonid Padyukov

Affiliation Rheumatology Unit, Department of Medicine, Karolinska Institutet and Karolinska University Hospital Solna, Stockholm, Sweden

Lars Alfredsson

Affiliation Institute of Environmental Medicine, Karolinska Institutet, Stockholm, Sweden

Lars Klareskog

Affiliation Rheumatology Unit, Department of Medicine, Karolinska Institutet and Karolinska University Hospital Solna, Stockholm, Sweden

Bengt Glimelius

Affiliations Department of Pathology and Oncology, Karolinska Institutet, Stockholm, Sweden, Department of Radiology, Oncology, and Radiation Sciences, Uppsala University, Uppsala, Sweden

Mads Melbye

Affiliation Department of Epidemiology Research, Statens Serum Institut, Copenhagen, Denmark

Edison T. Liu

Affiliation Human Genetics, Genome Institute of Singapore, A*STAR, Singapore, Singapore

Hans-Olov Adami

Affiliations Department of Medical Epidemiology and Biostatistics, Karolinska Institutet, Stockholm, Sweden, Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America

Keith Humphreys

These authors also contributed equally to this work.

Affiliation Department of Medical Epidemiology and Biostatistics, Karolinska Institutet, Stockholm, Sweden

Jianjun Liu

liuj3@gis.a-star.edu.sg

These authors also contributed equally to this work.

Affiliation Human Genetics, Genome Institute of Singapore, A*STAR, Singapore, Singapore

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KES ETL HOA JL. Performed the experiments: IDI. Analyzed the data: KES JNF CFS HD LC VK ER KH JL. Contributed reagents/materials/analysis tools: KES CFS LC HH ETC NR JRC LPR PNB PMB LA JR WC SD PH LMM RKS SSW SLS ZSF AJN NEK TMH BA AK SM MPP CMV PB QL SHZ YZ TZ SL JJS MTS SJC LP LA LK BG MM ETL HOA JL. Wrote the paper: KES JNF KH JL. Critical revision of manuscript: CFS HD LC VK ETC IDI SJC. Genotyped samples and provided data from the BC study: ARBW. Provided data from the SF1 and SF2 GWAS studies: JR. Supervised the experiments: JL JNF.