About the Authors

Naomi R. Wray

Naomi.Wray@qimr.edu.au

Affiliation Genetic Epidemiology and Queensland Statistical Genetics, Queensland Institute of Medical Research, Brisbane, Australia

Jian Yang

Affiliation Genetic Epidemiology and Queensland Statistical Genetics, Queensland Institute of Medical Research, Brisbane, Australia

Michael E. Goddard

Affiliations Department of Food and Agricultural Systems, University of Melbourne, Melbourne, Australia, Victoria Department of Primary Industries, Melbourne, Australia

Peter M. Visscher

Affiliation Genetic Epidemiology and Queensland Statistical Genetics, Queensland Institute of Medical Research, Brisbane, Australia

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NRW. Performed the experiments: NRW. Analyzed the data: NRW. Contributed reagents/materials/analysis tools: NRW JY MEG PMV. Wrote the paper: NRW MEG PMV.