About the Authors

Clive J. Hoggart

c.hoggart@ic.ac.uk

Affiliation Department of Epidemiology and Public Health, Imperial College, London, United Kingdom

John C. Whittaker

Affiliation Non-Communicable Disease Epidemiology Unit, London School of Hygiene and Tropical Medicine, London, United Kingdom

Maria De Iorio

Affiliation Department of Epidemiology and Public Health, Imperial College, London, United Kingdom

David J. Balding

Affiliation Department of Epidemiology and Public Health, Imperial College, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CJH JCW MDI DJB. Performed the experiments: CJH. Analyzed the data: CJH. Wrote the paper: CJH DJB.