About the Authors

Robert B. Weiss

Bob.Weiss@genetics.utah.edu (RBW); Mark.Leppert@genetics.utah.edu (MFL)

Affiliation Department of Human Genetics, Eccles Institute of Human Genetics, University of Utah School of Medicine, Salt Lake City, Utah, United States of America

Timothy B. Baker

Affiliation Department of Medicine, Center for Tobacco Research and Intervention, University of Wisconsin School of Medicine, Madison, Wisconsin, United States of America

Dale S. Cannon

Affiliation Department of Psychiatry, University of Utah School of Medicine, Salt Lake City, Utah, United States of America

Andrew von Niederhausern

Affiliation Department of Human Genetics, Eccles Institute of Human Genetics, University of Utah School of Medicine, Salt Lake City, Utah, United States of America

Diane M. Dunn

Affiliation Department of Human Genetics, Eccles Institute of Human Genetics, University of Utah School of Medicine, Salt Lake City, Utah, United States of America

Nori Matsunami

Affiliation Department of Human Genetics, Eccles Institute of Human Genetics, University of Utah School of Medicine, Salt Lake City, Utah, United States of America

Nanda A. Singh

Affiliation Department of Human Genetics, Eccles Institute of Human Genetics, University of Utah School of Medicine, Salt Lake City, Utah, United States of America

Lisa Baird

Affiliation Department of Human Genetics, Eccles Institute of Human Genetics, University of Utah School of Medicine, Salt Lake City, Utah, United States of America

Hilary Coon

Affiliation Department of Psychiatry, University of Utah School of Medicine, Salt Lake City, Utah, United States of America

William M. McMahon

Affiliation Department of Psychiatry, University of Utah School of Medicine, Salt Lake City, Utah, United States of America

Megan E. Piper

Affiliation Department of Medicine, Center for Tobacco Research and Intervention, University of Wisconsin School of Medicine, Madison, Wisconsin, United States of America

Michael C. Fiore

Affiliation Department of Medicine, Center for Tobacco Research and Intervention, University of Wisconsin School of Medicine, Madison, Wisconsin, United States of America

Mary Beth Scholand

Affiliation Department of Human Genetics, Eccles Institute of Human Genetics, University of Utah School of Medicine, Salt Lake City, Utah, United States of America

John E. Connett

Affiliation Division of Biostatistics, School of Public Health, University of Minnesota, Minneapolis, Minnesota, United States of America

Richard E. Kanner

Affiliation Department of Internal Medicine, Division of Respiratory, Critical Care and Occupational Pulmonary Medicine, University of Utah School of Medicine, Salt Lake City, Utah, United States of America

Lorise C. Gahring

Affiliations Department of Internal Medicine, Division of Respiratory, Critical Care and Occupational Pulmonary Medicine, University of Utah School of Medicine, Salt Lake City, Utah, United States of America, Geriatric Research Education and Clinical Center, Salt Lake City VA Medical Center, Salt Lake City, Utah, United States of America

Scott W. Rogers

Affiliations Geriatric Research Education and Clinical Center, Salt Lake City VA Medical Center, Salt Lake City, Utah, United States of America, Department of Neurobiology and Anatomy, University of Utah School of Medicine, Salt Lake City, Utah, United States of America

John R. Hoidal

Affiliation Department of Internal Medicine, Division of Respiratory, Critical Care and Occupational Pulmonary Medicine, University of Utah School of Medicine, Salt Lake City, Utah, United States of America

Mark F. Leppert

Bob.Weiss@genetics.utah.edu (RBW); Mark.Leppert@genetics.utah.edu (MFL)

Affiliation Department of Human Genetics, Eccles Institute of Human Genetics, University of Utah School of Medicine, Salt Lake City, Utah, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RBW TBB DSC AvN DMD NM NAS HC WMM MCF MBS LCG SWR JRH MFL. Performed the experiments: RBW TBB DSC AvN DMD NM LB MFL. Analyzed the data: RBW TBB DSC AvN DMD NM NAS LB HC WMM MEP MBS LCG SWR JRH MFL. Contributed reagents/materials/analysis tools: RBW TBB DSC NM LB MEP MCF JEC REK JRH MFL. Wrote the paper: RBW TBB DSC JRH MFL.