About the Authors

Brendan A Gavin

Affiliation Department of Biology, Dartmouth College, Hanover, New Hampshire, United States of America

Susan E Arruda

Affiliation Department of Biology, Franklin Pierce College, Rindge, New Hampshire, United States of America

Patrick J Dolph

To whom correspondence should be addressed. E-mail: patrick.dolph@dartmouth.edu

Affiliation Department of Biology, Dartmouth College, Hanover, New Hampshire, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

BAG, SEA, and PJD conceived and designed the experiments. BAG performed the experiments. BAG and PJD analyzed the data. BAG wrote the paper.