About the Authors

Luke E Berchowitz

Affiliations Department of Biology, The University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America , The Carolina Center for Genome Sciences, The University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Kirk E Francis

Affiliations Department of Biology, The University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America , The Carolina Center for Genome Sciences, The University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Alexandra L Bey

Affiliations Department of Biology, The University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America , The Carolina Center for Genome Sciences, The University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Gregory P Copenhaver

To whom correspondence should be addressed. E-mail: gcopenhaver@bio.unc.edu

Affiliations Department of Biology, The University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America , The Carolina Center for Genome Sciences, The University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

LEB, KEF, and GPC conceived and designed the experiments and wrote the paper. LEP, KEF, and ALB performed the experiments. LEB and GPC analyzed the data. GPC contributed reagents/materials/analysis tools.