About the Authors

John L Rinn

Affiliation Program in Epithelial Biology, Department of Dermatology, Stanford University School of Medicine, Stanford, California, United States of America

Chanda Bondre

Affiliation Department of Biochemistry, Stanford University School of Medicine, Stanford, California, United States of America

Hayes B Gladstone

Affiliation Program in Epithelial Biology, Department of Dermatology, Stanford University School of Medicine, Stanford, California, United States of America

Patrick O Brown

To whom correspondence should be addressed. E-mail: howchang@stanford.edu (HYC); pbrown@pmgm2.stanford.edu (POB)

Affiliations Department of Biochemistry, Stanford University School of Medicine, Stanford, California, United States of America , Howard Hughes Medical Institute, Stanford University School of Medicine, Stanford, California, United States of America

Howard Y Chang

To whom correspondence should be addressed. E-mail: howchang@stanford.edu (HYC); pbrown@pmgm2.stanford.edu (POB)

Affiliation Program in Epithelial Biology, Department of Dermatology, Stanford University School of Medicine, Stanford, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

JLR, POB, and HYC conceived and designed the experiments. JLR and CB performed the experiments. JLR and HYC analyzed the data. HBG and POB contributed reagents/materials/analysis tools. JLR, POB, and HYC wrote the paper.