About the Authors

Marcela Raices

Affiliation The Salk Institute for Biological Studies, La Jolla, California, United States of America

Hugo Maruyama

Affiliations The Salk Institute for Biological Studies, La Jolla, California, United States of America , Graduate School of Biostudies, Kyoto University, Kyoto, Japan

Andrew Dillin

Affiliation The Salk Institute for Biological Studies, La Jolla, California, United States of America

Jan Karlseder

To whom correspondence should be addressed. E-mail: karlseder@salk.edu

Affiliation The Salk Institute for Biological Studies, La Jolla, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

MR, AD, and JK conceived and designed the experiments. MR and HM performed the experiments. MR, HM, AD, and JK analyzed the data. AD contributed reagents/materials/analysis tools. AD and JK wrote the paper.