About the Authors

James M. Osborne

Contributed equally to this work with: James M. Osborne, Alexander G. Fletcher

jmosborne@unimelb.edu.au (JMO); a.g.fletcher@sheffield.ac.uk (AGF)

Affiliation School of Mathematics and Statistics, University of Melbourne, Parkville, Victoria, Australia

Alexander G. Fletcher

Contributed equally to this work with: James M. Osborne, Alexander G. Fletcher

jmosborne@unimelb.edu.au (JMO); a.g.fletcher@sheffield.ac.uk (AGF)

Affiliations School of Mathematics and Statistics, University of Sheffield, Sheffield, United Kingdom, Bateson Centre, University of Sheffield, Sheffield, United Kingdom

Joe M. Pitt-Francis

Affiliation Department of Computer Science, University of Oxford, Oxford, United Kingdom

Philip K. Maini

Affiliation Wolfson Centre for Mathematical Biology, Mathematical Institute, University of Oxford, Oxford, United Kingdom

David J. Gavaghan

Affiliation Department of Computer Science, University of Oxford, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: JMO AGF. Data curation: JMO AGF JMPF. Formal analysis: JMO AGF. Funding acquisition: PKM DJG. Investigation: JMO AGF. Methodology: JMO AGF. Project administration: JMO AGF. Resources: PKM DJG. Software: JMO AGF JMPF. Supervision: PKM DJG. Validation: JMO AGF JMPF. Visualization: JMO AGF JMPF. Writing – original draft: JMO AGF. Writing – review & editing: JMO AGF JMPF PKM DJG.