About the Authors

Yu Toyoshima

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Terumasa Tokunaga

Affiliations Department of Systems Design and Informatics, Faculty of Computer Science and Systems Engineering, Kyushu Institute of Technology, Iizuka-shi, Fukuoka, Japan, The Institute of Statistical Mathematics, Research Organization of Information and Systems, Tachikawa, Tokyo, Japan

Osamu Hirose

Affiliation Faculty of Electrical and Computer Engineering, Institute of Science and Engineering, Kanazawa University, Kakuma, Kanazawa, Japan

Manami Kanamori

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Takayuki Teramoto

Affiliation Department of Biology, Faculty of Sciences, Kyushu University, Higashi-ku, Fukuoka, Japan

Moon Sun Jang

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Sayuri Kuge

Affiliation Department of Biology, Faculty of Sciences, Kyushu University, Higashi-ku, Fukuoka, Japan

Takeshi Ishihara

Affiliation Department of Biology, Faculty of Sciences, Kyushu University, Higashi-ku, Fukuoka, Japan

Ryo Yoshida

Affiliation The Institute of Statistical Mathematics, Research Organization of Information and Systems, Tachikawa, Tokyo, Japan

Yuichi Iino

iino@bs.s.u-tokyo.ac.jp

Affiliations Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan, CREST, Japan Science and Technology Corporation, Bunkyo-ku, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Performed the experiments: MK MSJ TTe YT. Analyzed the data: YT YI TTo OH RY TTe MK MSJ SK TI. Wrote the paper: YT YI. Conceived the project: YT YI. Developed the method of image anlaysis: YT YI. Supported the development: TTo OH RY MK TI. Developed the customized microscope: TTe TI. Designed the experiments: YT YI TTe MK MSJ SK TI.