About the Authors

Patrick Gemmell

Affiliation Department of Zoology, University of Oxford, Oxford, United Kingdom

Jotun Hein

Affiliation Department of Statistics, University of Oxford, Oxford, United Kingdom

Aris Katzourakis

aris.katzourakis@zoo.ox.ac.uk

Affiliation Department of Zoology, University of Oxford, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Wrote the paper: PG JH AK. Conceived the study: PG JH AK. Developed and implemented the models: PG. Supervised development and implementation of models: JH AK.