About the Authors

Jerome Kelleher

jerome.kelleher@well.ox.ac.uk

Affiliation Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom

Alison M Etheridge

Affiliation Department of Statistics, University of Oxford, Oxford, United Kingdom

Gilean McVean

Affiliations Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom, Department of Statistics, University of Oxford, Oxford, United Kingdom, Li Ka Shing Centre for Health Information and Discovery, University of Oxford, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JK AME GM. Performed the experiments: JK. Analyzed the data: JK. Wrote the paper: JK GM.