About the Authors

Richard A. Morton

Affiliation Department of Biology, McMaster University, Hamilton, Ontario, Canada

Jonathan R. Stone

jstoner@mcmaster.ca

Affiliations Department of Biology, McMaster University, Hamilton, Ontario, Canada, Origins Institute, McMaster University, Hamilton, Ontario, Canada

Rama S. Singh

Affiliation Department of Biology, McMaster University, Hamilton, Ontario, Canada

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Wrote the paper: JRS RAM RSS. Conceived the study: RSS. Designed the computational model: RAM JRS. Encoded the computer program, ran simulations, and analyzed the data: RAM.