About the Authors

William R. Holmes

wrholmes@math.ubc.ca

Affiliation Department of Mathematics, University of British Columbia, Vancouver, British Columbia, Canada

Benjamin Lin

Affiliation Biomedical Engineering, Johns Hopkins University, Baltimore, Maryland, United States of America

Andre Levchenko

Affiliation Biomedical Engineering, Johns Hopkins University, Baltimore, Maryland, United States of America

Leah Edelstein-Keshet

Affiliation Department of Mathematics, University of British Columbia, Vancouver, British Columbia, Canada

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Wrote the paper: WRH BL AL LEK. Performed the modelling work: WRH LEK Performed associated experimental work: BL AL.