About the Authors

Sarah E. Hadyniak

Roles Conceptualization, Investigation, Methodology, Writing – original draft, Writing – review & editing

Affiliation Department of Biology, Johns Hopkins University, Baltimore, Maryland, United States

Joanna F. D. Hagen

Contributed equally to this work with: Joanna F. D. Hagen, Kiara C. Eldred

Roles Investigation, Methodology

Affiliation Department of Biology, Johns Hopkins University, Baltimore, Maryland, United States

Kiara C. Eldred

Contributed equally to this work with: Joanna F. D. Hagen, Kiara C. Eldred

Roles Conceptualization, Investigation, Methodology, Writing – review & editing

Affiliations Department of Biology, Johns Hopkins University, Baltimore, Maryland, United States, Department of Biological Structure, University of Washington, Seattle, Washington State, United States

Boris Brenerman

Roles Data curation, Formal analysis

Affiliation Department of Biology, Johns Hopkins University, Baltimore, Maryland, United States

Katarzyna A. Hussey

Roles Investigation, Methodology

Affiliation Department of Biology, Johns Hopkins University, Baltimore, Maryland, United States

Rajiv C. McCoy

Roles Data curation, Investigation

Affiliation Department of Biology, Johns Hopkins University, Baltimore, Maryland, United States

Michael E. G. Sauria

Roles Data curation

Affiliation Department of Biology, Johns Hopkins University, Baltimore, Maryland, United States

James A. Kuchenbecker

Roles Investigation

Affiliation Department of Ophthalmology, University of Washington, Seattle, Washington State, United States

Thomas Reh

Roles Supervision

Affiliation Department of Biological Structure, University of Washington, Seattle, Washington State, United States

Ian Glass

Roles Methodology

Affiliation Department of Biological Structure, University of Washington, Seattle, Washington State, United States

Maureen Neitz

Roles Investigation, Methodology

Affiliation Department of Ophthalmology, University of Washington, Seattle, Washington State, United States

Jay Neitz

Roles Supervision

Affiliation Department of Ophthalmology, University of Washington, Seattle, Washington State, United States

James Taylor

† Deceased.

Roles Conceptualization, Data curation, Supervision

Affiliation Department of Biology, Johns Hopkins University, Baltimore, Maryland, United States

Robert J. Johnston Jr

Roles Conceptualization, Funding acquisition, Project administration, Supervision, Visualization, Writing – original draft, Writing – review & editing

robertjohnston@jhu.edu

Affiliation Department of Biology, Johns Hopkins University, Baltimore, Maryland, United States

Competing Interests

The authors have declared that no competing interests exist.