About the Authors

Xuejun Tian

Contributed equally to this work with: Xuejun Tian, Upasana Gala

Affiliation Life Sciences Institute and Innovation Center for Cell Biology, Zhejiang University, Hangzhou, China

Upasana Gala

Contributed equally to this work with: Xuejun Tian, Upasana Gala

Affiliation Program in Developmental Biology, Baylor College of Medicine, Houston, Texas, United States of America

Yongping Zhang

Affiliation Life Sciences Institute and Innovation Center for Cell Biology, Zhejiang University, Hangzhou, China

Weina Shang

Affiliation Life Sciences Institute and Innovation Center for Cell Biology, Zhejiang University, Hangzhou, China

Sonal Nagarkar Jaiswal

Affiliation Department of Molecular and Human Genetics, Baylor College of Medicine, Houston, Texas, United States of America

Alberto di Ronza

Affiliations Department of Molecular and Human Genetics, Baylor College of Medicine, Houston, Texas, United States of America, Jan and Dan Duncan Neurological Research Institute, Texas Children’s Hospital, Houston, Texas, United States of America

Manish Jaiswal

Affiliations Department of Molecular and Human Genetics, Baylor College of Medicine, Houston, Texas, United States of America, Howard Hughes Medical Institute, Baylor College of Medicine, Houston, Texas, United States of America

Shinya Yamamoto

Affiliations Program in Developmental Biology, Baylor College of Medicine, Houston, Texas, United States of America, Department of Molecular and Human Genetics, Baylor College of Medicine, Houston, Texas, United States of America, Jan and Dan Duncan Neurological Research Institute, Texas Children’s Hospital, Houston, Texas, United States of America

Hector Sandoval

Affiliations Department of Molecular and Human Genetics, Baylor College of Medicine, Houston, Texas, United States of America, Jan and Dan Duncan Neurological Research Institute, Texas Children’s Hospital, Houston, Texas, United States of America

Lita Duraine

Affiliation Howard Hughes Medical Institute, Baylor College of Medicine, Houston, Texas, United States of America

Marco Sardiello

Affiliations Program in Developmental Biology, Baylor College of Medicine, Houston, Texas, United States of America, Department of Molecular and Human Genetics, Baylor College of Medicine, Houston, Texas, United States of America, Jan and Dan Duncan Neurological Research Institute, Texas Children’s Hospital, Houston, Texas, United States of America

Roy V. Sillitoe

Affiliations Jan and Dan Duncan Neurological Research Institute, Texas Children’s Hospital, Houston, Texas, United States of America, Department of Pathology and Immunology, Department of Neuroscience, Baylor College of Medicine, Houston, Texas, United States of America

Kartik Venkatachalam

Affiliations Program in Developmental Biology, Baylor College of Medicine, Houston, Texas, United States of America, Department of Integrative Biology and Pharmacology, University of Texas School of Medicine, Houston, Texas, United States of America

Hengyu Fan

Affiliation Life Sciences Institute and Innovation Center for Cell Biology, Zhejiang University, Hangzhou, China

Hugo J. Bellen

ctong@zju.edu.cn (CT), hbellen@bcm.edu (HJB)

Affiliations Program in Developmental Biology, Baylor College of Medicine, Houston, Texas, United States of America, Department of Molecular and Human Genetics, Baylor College of Medicine, Houston, Texas, United States of America, Jan and Dan Duncan Neurological Research Institute, Texas Children’s Hospital, Houston, Texas, United States of America, Howard Hughes Medical Institute, Baylor College of Medicine, Houston, Texas, United States of America, Department of Neuroscience, Baylor College of Medicine, Houston, Texas, United States of America

Chao Tong

ctong@zju.edu.cn (CT), hbellen@bcm.edu (HJB)

Affiliation Life Sciences Institute and Innovation Center for Cell Biology, Zhejiang University, Hangzhou, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CT HJB XT UG. Performed the experiments: CT XT UG YZ WS SNJ RVS MJ SY HS LD KV HF AdR MS. Analyzed the data: CT HJB XT UG. Wrote the paper: CT HJB XT UG.