About the Authors

Tobias Bast

To whom correspondence should be addressed. E-mail: tobias.bast@nottingham.ac.uk

Affiliations Centre for Cognitive and Neural Systems (CCNS), School of Biomedical Sciences, University of Edinburgh, Edinburgh, United Kingdom , School of Psychology and Institute of Neuroscience, University of Nottingham, Nottingham, United Kingdom

Iain A Wilson

† Deceased.

Affiliation Centre for Cognitive and Neural Systems (CCNS), School of Biomedical Sciences, University of Edinburgh, Edinburgh, United Kingdom

Menno P Witter

Affiliations Centre for the Biology of Memory and Kavli Institute for Systems Neuroscience, Norwegian University of Science and Technology (NTNU), Trondheim, Norway , Department of Anatomy and Neuroscience, VU University medical center, Amsterdam, the Netherlands

Richard G. M Morris

Affiliation Centre for Cognitive and Neural Systems (CCNS), School of Biomedical Sciences, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

TB conceived the overall design of the study, took main responsibility for the behavioral and field-potential experiments, performed all lesions and histological analysis, statistically analyzed the results, and wrote the paper; he also raised funding for the work. IAW took main responsibility for the single-unit experiments, including their design and recording-data analysis, and co-wrote the paper. MPW advised on the anatomical aspects of the work, contributed to the 3D brain reconstructions, and co-wrote the paper. RGMM contributed to the design of the study and co-wrote the paper; he also raised funding for the work.