About the Authors

Anja Forche

Affiliation Department of Genetics, Cell Biology and Development, University of Minnesota, Minneapolis, Minnesota, United States of America

Kevin Alby

Affiliation Department of Molecular Microbiology and Immunology, Brown University, Providence, Rhode Island, United States of America

Dana Schaefer

Affiliation Department of Molecular Microbiology and Immunology, Brown University, Providence, Rhode Island, United States of America

Alexander D Johnson

Affiliation Departments of Biochemistry and Biophysics and Microbiology and Immunology, University of California San Francisco, San Francisco, California, United States of America

Judith Berman

Affiliation Department of Genetics, Cell Biology and Development, University of Minnesota, Minneapolis, Minnesota, United States of America

Richard J Bennett

To whom correspondence should be addressed. E-mail: Richard_Bennett@brown.edu

Affiliation Department of Molecular Microbiology and Immunology, Brown University, Providence, Rhode Island, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

AF, ADJ, JB, and RJB conceived and designed the experiments. AF, KA, DS, and RJB performed the experiments. AF, KA, ADJ, JB, and RJB analyzed the data. AF, KA, ADJ, JB, and RJB wrote the paper.