About the Authors

Robert Illingworth

Affiliation Wellcome Trust Centre for Cell Biology, University of Edinburgh, Edinburgh, United Kingdom

Alastair Kerr

Affiliation Wellcome Trust Centre for Cell Biology, University of Edinburgh, Edinburgh, United Kingdom

Dina DeSousa

Affiliation Wellcome Trust Centre for Cell Biology, University of Edinburgh, Edinburgh, United Kingdom

Helle Jørgensen

Affiliation Lymphocyte Development Group, MRC Clinical Sciences Centre, Imperial College School of Medicine, London, United Kingdom

Peter Ellis

Affiliation Wellcome Trust Sanger Centre, Hinxton, Cambridge, United Kingdom

Jim Stalker

Affiliation Wellcome Trust Sanger Centre, Hinxton, Cambridge, United Kingdom

David Jackson

Affiliation Wellcome Trust Sanger Centre, Hinxton, Cambridge, United Kingdom

Chris Clee

Affiliation Wellcome Trust Sanger Centre, Hinxton, Cambridge, United Kingdom

Robert Plumb

Affiliation Wellcome Trust Sanger Centre, Hinxton, Cambridge, United Kingdom

Jane Rogers

Affiliation Wellcome Trust Sanger Centre, Hinxton, Cambridge, United Kingdom

Sean Humphray

Affiliation Wellcome Trust Sanger Centre, Hinxton, Cambridge, United Kingdom

Tony Cox

Affiliation Wellcome Trust Sanger Centre, Hinxton, Cambridge, United Kingdom

Cordelia Langford

Affiliation Wellcome Trust Sanger Centre, Hinxton, Cambridge, United Kingdom

Adrian Bird

To whom correspondence should be addressed. E-mail: a.bird@ed.ac.uk

Affiliation Wellcome Trust Centre for Cell Biology, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

AB, RI, and HJ conceived and designed the experiments. RI and DD performed the experiments. RI, AK, JS, DJ, and TC analyzed the data. HJ contributed reagents/materials/analysis tools. PE, SH, and CL designed and printed the CGI microarrays. CC, RP, and JR performed DNA sequencing of the CGI library. RI and AB wrote the paper.