About the Authors

Tessa M Burch-Smith

Affiliation Department of Molecular, Cellular, and Developmental Biology, Yale University, New Haven, Connecticut, United States of America

Michael Schiff

¤ Current address: Ceres, Thousand Oaks, California, United States of America

Affiliation Department of Molecular, Cellular, and Developmental Biology, Yale University, New Haven, Connecticut, United States of America

Jeffrey L Caplan

Affiliation Department of Molecular, Cellular, and Developmental Biology, Yale University, New Haven, Connecticut, United States of America

Jeffrey Tsao

Affiliation Department of Molecular, Cellular, and Developmental Biology, Yale University, New Haven, Connecticut, United States of America

Kirk Czymmek

Affiliation Department of Biological Sciences, Delaware Biotechnology Institute, University of Delaware, Newark, Delaware, United States of America

Savithramma P Dinesh-Kumar

To whom correspondence should be addressed. E-mail: savithramma.dinesh-kumar@yale.edu

Affiliation Department of Molecular, Cellular, and Developmental Biology, Yale University, New Haven, Connecticut, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

TMB and SPD conceived and designed the experiments and wrote the paper. TMB, MS, JLC, JT, and KC performed the experiments. TMB, JLC, JT, KC, and SPD analyzed the data. TMB, MS, and KC contributed reagents/materials/analysis tools.