About the Authors

Sagiv Shifman

Affiliation Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom

Jordana Tzenova Bell

Affiliation Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom

Richard R Copley

Affiliation Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom

Martin S Taylor

Affiliation Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom

Robert W Williams

Affiliation Department of Anatomy and Neurobiology, University of Tennessee Health Science Center, Memphis, Tennessee, United States of America

Richard Mott

Affiliation Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom

Jonathan Flint

To whom correspondence should be addressed. E-mail: jf@well.ox.ac.uk

Affiliation Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

RWW, RM, and JF conceived and designed the experiments. SS, JTB RRC, MST, and RM analyzed the data. RWW and RM contributed reagents/materials/analysis tools. SS, JTB, and JF wrote the paper. SS and JTB performed data cleaning and statistical analysis. RRC and MST performed the sequence analysis. RW coordinated sample collection. RM and JF formulated and directed the project.