About the Authors

Ping Hou

Affiliations Department of Medicine and Section of Rheumatology, University of Chicago, Chicago, Illinois, United States of America , Committee on Immunology, University of Chicago, Chicago, Illinois, United States of America

Elizabeth Araujo

Affiliations Department of Medicine and Section of Rheumatology, University of Chicago, Chicago, Illinois, United States of America , Committee on Immunology, University of Chicago, Chicago, Illinois, United States of America

Tong Zhao

Affiliations Department of Chemistry, University of Chicago, Chicago, Illinois, United States of America , Institute for Biophysical Dynamics, University of Chicago, Chicago, Illinois, United States of America , James Franck Institute, University of Chicago, Chicago, Illinois, United States of America

Miao Zhang

Affiliation Department of Medicine and Section of Rheumatology, University of Chicago, Chicago, Illinois, United States of America

Don Massenburg

Affiliations Department of Medicine and Section of Rheumatology, University of Chicago, Chicago, Illinois, United States of America , Committee on Immunology, University of Chicago, Chicago, Illinois, United States of America

Margaret Veselits

Affiliations Department of Medicine and Section of Rheumatology, University of Chicago, Chicago, Illinois, United States of America , Committee on Immunology, University of Chicago, Chicago, Illinois, United States of America

Colleen Doyle

Affiliation Committee on Immunology, University of Chicago, Chicago, Illinois, United States of America

Aaron R Dinner

Affiliations Committee on Immunology, University of Chicago, Chicago, Illinois, United States of America , Department of Chemistry, University of Chicago, Chicago, Illinois, United States of America , Institute for Biophysical Dynamics, University of Chicago, Chicago, Illinois, United States of America , James Franck Institute, University of Chicago, Chicago, Illinois, United States of America

Marcus R Clark

To whom correspondence should be addressed. E-mail: mclark@medicine.bsd.uchicago.edu

Affiliations Department of Medicine and Section of Rheumatology, University of Chicago, Chicago, Illinois, United States of America , Committee on Immunology, University of Chicago, Chicago, Illinois, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

MRC conceived and designed the experiments. PH, EA, and MV performed the experiments. TZ, ARD, and MRC analyzed the data. MZ, DM, and CD contributed reagents/materials/analysis tools. ARD and MRC wrote the paper.