About the Authors

Benoît Goossens

Contributed equally to this work with: Benoît Goossens, Lounès Chikhi

To whom correspondence should be addressed. E-mail: goossensbr@cardiff.ac.uk (BG), Email: chikhi@cict.fr (LC)

Affiliations Biodiversity and Ecological Processes Group, Cardiff School of Biosciences, Cardiff University, Cardiff, United Kingdom, UMR 5174 Evolution et Diversité Biologique, Université Paul Sabatier, Toulouse, France, Kinabatangan Orang-utan Conservation Project, Sandakan, Sabah, Malaysia, ITBC, Universiti Malaysia Sabah, Kota Kinabalu, Sabah, Malaysia

Lounès Chikhi

Contributed equally to this work with: Benoît Goossens, Lounès Chikhi

To whom correspondence should be addressed. E-mail: goossensbr@cardiff.ac.uk (BG), Email: chikhi@cict.fr (LC)

Affiliation UMR 5174 Evolution et Diversité Biologique, Université Paul Sabatier, Toulouse, France

Marc Ancrenaz

Affiliation Kinabatangan Orang-utan Conservation Project, Sandakan, Sabah, Malaysia

Isabelle Lackman-Ancrenaz

Affiliations Kinabatangan Orang-utan Conservation Project, Sandakan, Sabah, Malaysia, Pittsburgh Zoo, Pittsburgh, Pennsylvania, United States of America

Patrick Andau

Affiliation Sabah Wildlife Department, Wisma Muis, Kota Kinabalu, Sabah, Malaysia

Michael W Bruford

Affiliation Biodiversity and Ecological Processes Group, Cardiff School of Biosciences, Cardiff University, Cardiff, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

BG, LC, MA, ILA, PA, and MWB conceived and designed the experiments. BG and LC performed the experiments. BG, LC, and MA analyzed the data. MA, ILA, PA, and MWB contributed reagents/materials/analysis tools. BG, LC, and MWB wrote the paper.