About the Authors

William Mair

Affiliation Centre for Research on Ageing, University College London, Department of Biology, London, United Kingdom

Matthew D. W Piper

Affiliation Centre for Research on Ageing, University College London, Department of Biology, London, United Kingdom

Linda Partridge

To whom correspondence should be addressed. E-mail: l.partridge@ucl.ac.uk

Affiliation Centre for Research on Ageing, University College London, Department of Biology, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

LP and WM conceived and designed the experiments. WM performed the experiments and analyzed the data. LP, WM, and MDWP wrote the paper.