About the Authors

Emma T. Cadman

Affiliation The Royal Veterinary College, Department of Comparative Biomedical Sciences, London, United Kingdom

Katherine A. Thysse

Affiliation The Royal Veterinary College, Department of Comparative Biomedical Sciences, London, United Kingdom

Siobhan Bearder

Affiliation The Royal Veterinary College, Department of Comparative Biomedical Sciences, London, United Kingdom

Anita Y. N. Cheung

Affiliation The Royal Veterinary College, Department of Comparative Biomedical Sciences, London, United Kingdom

Ashleigh C. Johnston

Affiliation The Royal Veterinary College, Department of Comparative Biomedical Sciences, London, United Kingdom

James J. Lee

Affiliation Division of Pulmonary Medicine, Department of Biochemistry and Molecular Biology, Mayo Clinic Arizona, Scottsdale, Arizona, United States of America

Rachel A. Lawrence

rlawrence@rvc.ac.uk

Affiliation The Royal Veterinary College, Department of Comparative Biomedical Sciences, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ETC RAL. Performed the experiments: ETC KAT AYNC SB ACJ RAL. Analyzed the data: ETC KAT SB ACJ RAL. Contributed reagents/materials/analysis tools: JJL RAL. Wrote the paper: ETC RAL.